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The Carboniferous Portal

Introduction

The Carboniferous (/ˌkɑːrbəˈnɪfərəs/ KAR-bə-NIF-ər-əs) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period 358.9 Ma (million years ago) to the beginning of the Permian Period, 298.9 Ma. In North America, the Carboniferous is often treated as two separate geological periods, the earlier Mississippian and the later Pennsylvanian.

The name Carboniferous means "coal-bearing", from the Latin carbō ("coal") and ferō ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern "system" names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession.

Carboniferous is the period during which both terrestrial animal and land plant life was well established. Stegocephalia (four-limbed vertebrates including true tetrapods), whose forerunners (tetrapodomorphs) had evolved from lobe-finned fish during the preceding Devonian period, became pentadactylous during the Carboniferous. The period is sometimes called the Age of Amphibians because of the diversification of early amphibians such as the temnospondyls, which became dominant land vertebrates, as well as the first appearance of amniotes including synapsids (the clade to which modern mammals belong) and sauropsids (which include modern reptiles and birds) during the late Carboniferous. Due to the raised atmospheric oxygen level, land arthropods such as arachnids (e.g. trigonotarbids and Pulmonoscorpius), myriapods (e.g. Arthropleura) and insects (e.g. Meganeura) also underwent a major evolutionary radiation during the late Carboniferous. Vast swaths of forests and swamps covered the land, which eventually became the coal beds characteristic of the Carboniferous stratigraphy evident today.

The later half of the period experienced glaciations, low sea level, and mountain building as the continents collided to form Pangaea. A minor marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred at the end of the period, caused by climate change. (Full article...)

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The subphylum (or phylum) Chelicerata constitutes one of the major subdivisions of the phylum (or superphylum) Arthropoda, and includes horseshoe crabs, scorpions, spiders, mites, harvestmen, ticks, and Solifugae. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered in a cuticle made of chitin and proteins. The chelicerate bauplan consists of two tagmata, the cephalothorax and the abdomen. The group is named for their chelicerae, appendages near the mouth generally used to feed. The group has the open circulatory system typical of arthropods, in which a tube-like heart pumps blood through the hemocoel, which is the major body cavity.

Chelicerates were originally predators, but the group has diversified to use all the major feeding strategies. The guts of most modern chelicerates are too narrow for solid food, and they generally liquidize their food by grinding it with their chelicerae and pedipalps and flooding it with digestive enzymes. Most lay eggs that hatch as what look like miniature adults. In most chelicerate species the young have to fend for themselves, but in scorpions and some species of spider the females protect and feed their young.

The chelicerata originated as marine animals, possibly in the Cambrian period, but the first confirmed chelicerate fossils, eurypterids, date from 445 million years ago in the Late Ordovician period. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are chelicerates. (see more...)

Did you know...

... that the largest known ovules produced by any non-flowering seed-plant came from the Medullosales (fossilized leaves pictured), an order of extinct seed ferns?
... that Adolf Carl Noé challenged disbelief in the possibility of North American coal balls (example pictured) by presenting a wheelbarrow full of them?
... that the extinct griffenfly genus Bohemiatupus inhabited peat-mires?
... that Okmulgee State Park (pictured) in Oklahoma is one of only a few places in the world where the fossil coral Gymnophyllum wardi is found?
... that the type specimen of the extinct whip scorpion Graeophonus carbonarius was originally identified as a species of dragonfly?

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Photomicrograph of fossiliferous limestone (bisoparite or grainstone) from the Stoner Member of the Stanton Formation (Pennsylvanian) from the Ash Grove Quarry near Louisville, Nebraska. Coiled fossil is a fusilinid foraminifera. Coordinates are approximate.
Photo credit: Rygel, M.C.

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Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Cope distinguished himself as a child prodigy, publishing his first scientific paper at the age of nineteen. Cope later married and moved from Philadelphia to Haddonfield, New Jersey, although Cope would maintain a residence and museum in Philadelphia in his later years.

Cope had little formal scientific training, and he eschewed a teaching position for field work. He made regular trips to the American West prospecting in the 1870s and 1880s, often as a member of United States Geological Survey teams. A personal feud between Cope and paleontologist Othniel Charles Marsh led to a period of intense fossil-finding competition now known as the Bone Wars. Cope's financial fortunes soured after failed mining ventures in the 1880s. He experienced a resurgence in his career toward the end of his life before dying in 1897.

Cope's scientific pursuits nearly bankrupted him, but his contributions helped to define the field of American paleontology. He was a prodigious writer, with 1,400 papers published over his lifetime, although his rivals would debate the accuracy of his rapidly published works. He discovered, described, and named more than 1,000 vertebrate species including hundreds of fishes and dozens of dinosaurs. His proposals on the origin of mammalian molars and for the gradual enlargement of mammalian species over geologic time ("Cope's Law") are notable among his theoretical contributions. (see more...)