Marine invertebrates exhibit a wide range of modifications to survive in poorly oxygenated waters, including breathing tubes as in mollusc siphons. Fish have gills instead of lungs, although some species of fish, such as the lungfish, have both. Marine mammals (e.g. dolphins, whales, otters, and seals) need to surface periodically to breathe air. (Full article...)
Seals range in size from the 1 m (3 ft 3 in) and 45 kg (100 lb) Baikal seal to the 5 m (16 ft) and 3,200 kg (7,100 lb) southern elephant seal. Several species exhibit sexual dimorphism. They have streamlined bodies and four limbs that are modified into flippers. Though not as fast in the water as dolphins, seals are more flexible and agile. Otariids primarily use their front limbs to propel themselves through the water, while phocids and walruses primarily use their hind limbs for this purpose. Otariids and walruses have hind limbs that can be pulled under the body and used as legs on land. By comparison, terrestrial locomotion by phocids is more cumbersome. Otariids have visible external ears, while phocids and walruses lack these. Pinnipeds have well-developed senses—their eyesight and hearing are adapted for both air and water, and they have an advanced tactile system in their whiskers or vibrissae. Some species are well adapted for diving to great depths. They have a layer of fat, or blubber, under the skin to keep warm in cold water, and, other than the walrus, all species are covered in fur. (Full article...)
Image 2
3D reconstruction of the skull, viewed from the bottom left, exposing the teeth and internal skull roof
Cartorhynchus (meaning "shortened snout") is an extinct genus of earlyichthyosauriformmarine reptile that lived during the Early Triassicepoch, about 248 million years ago. The genus contains a single species, Cartorhynchus lenticarpus, named in 2014 by Ryosuke Motani and colleagues from a single nearly-complete skeleton found near Chaohu, Anhui Province, China. Along with its close relative Sclerocormus, Cartorhynchus was part of a diversification of marine reptiles that occurred suddenly (over about one million years) during the Spathiansubstage, soon after the devastating Permian-Triassic extinction event, but they were subsequently driven to extinction by volcanism and sea level changes by the Middle Triassic.
Measuring about 40 centimetres (16 in) long, Cartorhynchus was a small animal with a lizard-like body and a short torso; it probably swam in an eel-like manner at slow speeds. Its limbs bore extensive cartilage and could bend like flippers, which may have allowed it to walk on land. The most distinctive features of Cartorhynchus were its short, constricted snout, and its multiple rows of molar-like teeth which grew on the inside surface of its jaw bones. These teeth were not discovered until the specimen was subjected to CT scanning. Cartorhynchus likely preyed on hard-shelled invertebrates using suction feeding, although how it exactly used its inward-directed teeth is not yet known. It was one of up to five independent acquisitions of molar-like teeth among ichthyosauriforms. (Full article...)
Depending on the species, adult ctenophores range from a few millimeters to 1.5 m (5 ft) in size. Only 186 living species are currently recognised. (Full article...)
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The porbeagle or porbeagle shark (Lamna nasus) is a species of mackerel shark in the familyLamnidae, distributed widely in the cold and temperate marine waters of the North Atlantic and Southern Hemisphere. In the North Pacific, its ecological equivalent is the closely related salmon shark (L. ditropis). It typically reaches 2.5 m (8.2 ft) in length and a weight of 135 kg (298 lb); North Atlantic sharks grow larger than Southern Hemisphere sharks and differ in coloration and aspects of life history. Gray above and white below, the porbeagle has a very stout midsection that tapers towards the long, pointed snout and the narrow base of the tail. It has large pectoral and first dorsal fins, tiny pelvic, second dorsal, and anal fins, and a crescent-shaped caudal fin. The most distinctive features of this species are its three-cusped teeth, the white blotch at the aft base of its first dorsal fin, and the two pairs of lateral keels on its tail.
The porbeagle is an opportunistic hunter that preys mainly on bony fishes and cephalopods throughout the water column, including the bottom. Most commonly found over food-rich banks on the outer continental shelf, it makes occasional forays both close to shore and into the open ocean to a depth of 1,360 m (4,460 ft). It also conducts long-distance seasonal migrations, generally shifting between shallower and deeper water. The porbeagle is fast and highly active, with physiological adaptations that enable it to maintain a higher body temperature than the surrounding water. It can be solitary or gregarious, and has been known to perform seemingly playful behavior. This shark is aplacental viviparous with oophagy, developing embryos being retained within the mother's uterus and subsisting on non-viable eggs. Females typically bear four pups every year. (Full article...)
The oceanic whitetip shark (Carcharhinus longimanus) is a large pelagicrequiem shark inhabiting tropical and warm temperate seas. It has a stocky body with long, white-tipped, rounded fins. The species is typically solitary, though they may gather in large numbers at food concentrations. Bony fish and cephalopods are the main components of its diet and females give live birth.
Though slow-moving, it is opportunistic and aggressive, and is reputed to be dangerous to shipwreck survivors. The IUCN Red List considers the species to be critically endangered. Recent studies show steeply declining populations as they are harvested for their fins and meat. As with other shark species, the whitetip faces mounting fishing pressure throughout its range. (Full article...)
Sea urchins or urchins (/ˈɜːrtʃɪnz/) are typically spiny, globular animals, echinoderms in the class Echinoidea. About 950 species live on the seabed, inhabiting all oceans and depth zones from the intertidal to 5,000 metres (16,000 ft; 2,700 fathoms). Their tests (hard shells) are round and spiny, typically from 3 to 10 cm (1 to 4 in) across. Sea urchins move slowly, crawling with their tube feet, and sometimes pushing themselves with their spines. They feed primarily on algae but also eat slow-moving or sessile animals. Their predators include sea otters, starfish, wolf eels, and triggerfish.
Like all echinoderms, adult sea urchins have fivefold symmetry with their pluteus larvae featuring bilateral (mirror) symmetry; The latter indicates that they belong to the Bilateria, along with chordates, arthropods, annelids and molluscs. Sea urchins are found in every ocean and in every climate, from the tropics to the polar regions, and inhabit marine benthic (sea bed) habitats, from rocky shores to hadal zone depths. The fossil record of the Echinoids dates from the Ordovician period, some 450 million years ago. The closest echinoderm relatives of the sea urchin are the sea cucumbers (Holothuroidea), which like them are deuterostomes, a clade that includes the chordates. (Sand dollars are a separate order in the sea urchin class Echinoidea.) (Full article...)
A typical sea anemone is a single polyp attached to a hard surface by its base, but some species live in soft sediment, and a few float near the surface of the water. The polyp has a columnar trunk topped by an oral disc with a ring of tentacles and a central mouth. The tentacles can be retracted inside the body cavity or expanded to catch passing prey. They are armed with cnidocytes (stinging cells). In many species, additional nourishment comes from a symbiotic relationship with single-celleddinoflagellates, with zooxanthellae, or with green algae, zoochlorellae, that live within the cells. Some species of sea anemone live in association with clownfish, hermit crabs, small fish, or other animals to their mutual benefit. (Full article...)
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The dusky dolphin (Lagenorhynchus obscurus) is a dolphin found in coastal waters in the Southern Hemisphere. Its specific epithet is Latin for "dark" or "dim". It is very closely genetically related to the Pacific white-sided dolphin, but current scientific consensus holds they are distinct species. The dolphin's range is patchy, with major populations around South America, southwestern Africa, New Zealand, and various oceanic islands, with some sightings around southern Australia and Tasmania. The dusky dolphin prefers cool currents and inshore waters, but can also be found offshore. It feeds on a variety of fish and squid species and has flexible hunting tactics. The dusky dolphin is known for its remarkable acrobatics, having a number of aerial behaviours. The status of the dolphin is unknown, but it has been commonly caught in gill nets. (Full article...)
The Echiura, or spoon worms, are a small group of marineanimals. Once treated as a separate phylum, they are now considered to belong to Annelida. Annelids typically have their bodies divided into segments, but echiurans have secondarily lost their segmentation. The majority of echiurans live in burrows in soft sediment in shallow water, but some live in rock crevices or under boulders, and there are also deep sea forms. More than 230 species have been described. Spoon worms are cylindrical, soft-bodied animals usually possessing a non-retractable proboscis which can be rolled into a scoop-shape to feed. In some species the proboscis is ribbon-like, longer than the trunk and may have a forked tip. Spoon worms vary in size from less than a centimetre in length to more than a metre.
A tunicate is a marine invertebrate animal, a member of the subphylumTunicata (/ˌtjuːnɪˈkeɪtə/TEW-nih-KAY-tə). It is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates). The subphylum was at one time called Urochordata, and the term urochordates is still sometimes used for these animals. They are the only chordates that have lost their myomeric segmentation, with the possible exception of the seriation of the gill slits. However, doliolids still display segmentation of the muscle bands.
Some tunicates live as solitary individuals, but others replicate by budding and become colonies, each unit being known as a zooid. They are marine filter feeders with a water-filled, sac-like body structure and two tubular openings, known as siphons, through which they draw in and expel water. During their respiration and feeding, they take in water through the incurrent (or inhalant) siphon and expel the filtered water through the excurrent (or exhalant) siphon. Adult ascidian tunicates are sessile, immobile and permanently attached to rocks or other hard surfaces on the ocean floor. Thaliaceans (pyrosomes, doliolids, and salps) and larvaceans on the other hand, swim in the pelagic zone of the sea as adults. (Full article...)
Algae constitute a polyphyletic group since they do not include a common ancestor, and although their plastids seem to have a single origin, from cyanobacteria, they were acquired in different ways. Green algae are examples of algae that have primary chloroplasts derived from endosymbiotic cyanobacteria. Diatoms and brown algae are examples of algae with secondary chloroplasts derived from an endosymbiotic red alga. Algae exhibit a wide range of reproductive strategies, from simple asexual cell division to complex forms of sexual reproduction. (Full article...)
Image 19Cycling of marine phytoplankton. Phytoplankton live in the photic zone of the ocean, where photosynthesis is possible. During photosynthesis, they assimilate carbon dioxide and release oxygen. If solar radiation is too high, phytoplankton may fall victim to photodegradation. For growth, phytoplankton cells depend on nutrients, which enter the ocean by rivers, continental weathering, and glacial ice meltwater on the poles. Phytoplankton release dissolved organic carbon (DOC) into the ocean. Since phytoplankton are the basis of marine food webs, they serve as prey for zooplankton, fish larvae and other heterotrophic organisms. They can also be degraded by bacteria or by viral lysis. Although some phytoplankton cells, such as dinoflagellates, are able to migrate vertically, they are still incapable of actively moving against currents, so they slowly sink and ultimately fertilize the seafloor with dead cells and detritus. (from Marine food web)
Image 21A microbial mat encrusted with iron oxide on the flank of a seamount can harbour microbial communities dominated by the iron-oxidizing Zetaproteobacteria (from Marine prokaryotes)
Image 22Reconstruction of an ammonite, a highly successful early cephalopod that first appeared in the Devonian (about 400 mya). They became extinct during the same extinction event that killed the land dinosaurs (about 66 mya). (from Marine invertebrates)
Image 23Some representative ocean animal life (not drawn to scale) within their approximate depth-defined ecological habitats. Marine microorganisms exist on the surfaces and within the tissues and organs of the diverse life inhabiting the ocean, across all ocean habitats. (from Marine habitat)
Image 24Ocean surface chlorophyll concentrations in October 2019. The concentration of chlorophyll can be used as a proxy to indicate how many phytoplankton are present. Thus on this global map green indicates where a lot of phytoplankton are present, while blue indicates where few phytoplankton are present. – NASA Earth Observatory 2019. (from Marine food web)
Image 25
Different bacteria shapes (cocci, rods and spirochetes) and their sizes compared with the width of a human hair. A few bacteria are comma-shaped (vibrio). Archaea have similar shapes, though the archaeon Haloquadratum is flat and square.
The unit μm is a measurement of length, the micrometer, equal to 1/1,000 of a millimeter
Image 27Schematic representation of the changes in abundance between trophic groups in a temperate rocky reef ecosystem. (a) Interactions at equilibrium. (b) Trophic cascade following disturbance. In this case, the otter is the dominant predator and the macroalgae are kelp. Arrows with positive (green, +) signs indicate positive effects on abundance while those with negative (red, -) indicate negative effects on abundance. The size of the bubbles represents the change in population abundance and associated altered interaction strength following disturbance. (from Marine food web)
Image 28Scanning electron micrograph of a strain of Roseobacter, a widespread and important genus of marine bacteria. For scale, the membrane pore size is 0.2 μm in diameter. (from Marine prokaryotes)
Image 31Marine Species Changes in Latitude and Depth in three different ocean regions(1973–2019) (from Marine food web)
Image 32Topological positions versus mobility: (A) bottom-up groups (sessile and drifters), (B) groups at the top of the food web. Phyto, phytoplankton; MacroAlga, macroalgae; Proto, pelagic protozoa; Crus, Crustacea; PelBact, pelagic bacteria; Echino, Echinoderms; Amph, Amphipods; HerbFish, herbivorous fish; Zoopl, zooplankton; SuspFeed, suspension feeders; Polych, polychaetes; Mugil, Mugilidae; Gastropod, gastropods; Blenny, omnivorous blennies; Decapod, decapods; Dpunt, Diplodus puntazzo; Macropl, macroplankton; PlFish, planktivorous fish; Cephalopod, cephalopods; Mcarni, macrocarnivorous fish; Pisc, piscivorous fish; Bird, seabirds; InvFeed1 through InvFeed4, benthic invertebrate feeders. (from Marine food web)
Image 33
Diagram of a mycoloop (fungus loop)
Parasitic chytrids can transfer material from large inedible phytoplankton to zooplankton. Chytrids zoospores are excellent food for zooplankton in terms of size (2–5 μm in diameter), shape, nutritional quality (rich in polyunsaturated fatty acids and cholesterols). Large colonies of host phytoplankton may also be fragmented by chytrid infections and become edible to zooplankton. (from Marine fungi)
Image 34In the open ocean, sunlit surface epipelagic waters get enough light for photosynthesis, but there are often not enough nutrients. As a result, large areas contain little life apart from migrating animals. (from Marine habitat)
Image 35Lampreys are often parasitic and have a toothed, funnel-like sucking mouth (from Marine vertebrate)
Image 44Conceptual diagram of faunal community structure and food-web patterns along fluid-flux gradients within Guaymas seep and vent ecosystems. (from Marine food web)
Image 52An in situ perspective of a deep pelagic food web derived from ROV-based observations of feeding, as represented by 20 broad taxonomic groupings. The linkages between predator to prey are coloured according to predator group origin, and loops indicate within-group feeding. The thickness of the lines or edges connecting food web components is scaled to the log of the number of unique ROV feeding observations across the years 1991–2016 between the two groups of animals. The different groups have eight colour-coded types according to main animal types as indicated by the legend and defined here: red, cephalopods; orange, crustaceans; light green, fish; dark green, medusa; purple, siphonophores; blue, ctenophores and grey, all other animals. In this plot, the vertical axis does not correspond to trophic level, because this metric is not readily estimated for all members. (from Marine food web)
Image 53Diagram above contains clickable links
Image 54Chytrid parasites of marine diatoms. (A) Chytrid sporangia on Pleurosigma sp. The white arrow indicates the operculate discharge pore. (B) Rhizoids (white arrow) extending into diatom host. (C) Chlorophyll aggregates localized to infection sites (white arrows). (D and E) Single hosts bearing multiple zoosporangia at different stages of development. The white arrow in panel E highlights branching rhizoids. (F) Endobiotic chytrid-like sporangia within diatom frustule. Bars = 10 μm. (from Marine fungi)
Image 55The Ocean Cleanup is one of many organizations working toward marine conservation such at this interceptor vessel that prevents plastic from entering the ocean. (from Marine conservation)
Image 56The deep sea amphipodEurythenes plasticus, named after microplastics found in its body, demonstrating plastic pollution affects marine habitats even 6000m below sea level. (from Marine habitat)
Image 57Phylogenetic and symbiogenetic tree of living organisms, showing a view of the origins of eukaryotes and prokaryotes (from Marine fungi)
Image 58Coral reefs provide marine habitats for tube sponges, which in turn become marine habitats for fishes (from Marine habitat)
Image 59The distribution of anthropogenic stressors faced by marine species threatened with extinction in various marine regions of the world. Numbers in the pie charts indicate the percentage contribution of an anthropogenic stressors' impact in a specific marine region. (from Marine food web)
Image 60Sandy shores provide shifting homes to many species (from Marine habitat)
Image 63Conference events, such as the events hosted by the United Nations, help to bring together many stakeholders for awareness and action. (from Marine conservation)
Image 64This algae bloom occupies sunlit epipelagic waters off the southern coast of England. The algae are maybe feeding on nutrients from land runoff or upwellings at the edge of the continental shelf. (from Marine habitat)
Image 70Common-enemy graph of Antarctic food web. Potter Cove 2018. Nodes represent basal species and links indirect interactions (shared predators). Node and link widths are proportional to number of shared predators. Node colors represent functional groups. (from Marine food web)
Image 71
The global continental shelf, highlighted in light green, defines the extent of marine coastal habitats, and occupies 5% of the total world area
Image 72Elevation-area graph showing the proportion of land area at given heights and the proportion of ocean area at given depths (from Marine habitat)
Image 73Only 29 percent of the world surface is land. The rest is ocean, home to the marine habitats. The oceans are nearly four kilometres deep on average and are fringed with coastlines that run for nearly 380,000 kilometres.
Image 75Jellyfish are easy to capture and digest and may be more important as food sources than was previously thought. (from Marine food web)
Image 76Biomass pyramids. Compared to terrestrial biomass pyramids, aquatic pyramids are generally inverted at the base. (from Marine food web)
Image 77The pelagic food web, showing the central involvement of marine microorganisms in how the ocean imports nutrients from and then exports them back to the atmosphere and ocean floor (from Marine food web)
Image 80Anthropogenic stressors to marine species threatened with extinction (from Marine food web)
Image 81Estuaries occur when rivers flow into a coastal bay or inlet. They are nutrient rich and have a transition zone which moves from freshwater to saltwater. (from Marine habitat)
Model of the energy generating mechanism in marine bacteria
(1) When sunlight strikes a rhodopsin molecule (2) it changes its configuration so a proton is expelled from the cell (3) the chemical potential causes the proton to flow back to the cell (4) thus generating energy (5) in the form of adenosine triphosphate. (from Marine prokaryotes)
Image 85Food web structure in the euphotic zone. The linear food chain large phytoplankton-herbivore-predator (on the left with red arrow connections) has fewer levels than one with small phytoplankton at the base. The microbial loop refers to the flow from the dissolved organic carbon (DOC) via heterotrophic bacteria (Het. Bac.) and microzooplankton to predatory zooplankton (on the right with black solid arrows). Viruses play a major role in the mortality of phytoplankton and heterotrophic bacteria, and recycle organic carbon back to the DOC pool. Other sources of dissolved organic carbon (also dashed black arrows) includes exudation, sloppy feeding, etc. Particulate detritus pools and fluxes are not shown for simplicity. (from Marine food web)
Image 86A 2016 metagenomic representation of the tree of life using ribosomal protein sequences. The tree includes 92 named bacterial phyla, 26 archaeal phyla and five eukaryotic supergroups. Major lineages are assigned arbitrary colours and named in italics with well-characterized lineage names. Lineages lacking an isolated representative are highlighted with non-italicized names and red dots. (from Marine prokaryotes)
Image 88Ocean or marine biomass, in a reversal of terrestrial biomass, can increase at higher trophic levels. (from Marine food web)
Image 89On average there are more than one million microbial cells in every drop of seawater, and their collective metabolisms not only recycle nutrients that can then be used by larger organisms but also catalyze key chemical transformations that maintain Earth's habitability. (from Marine food web)
Image 90Phylogenetic tree representing bacterial OTUs from clone libraries and next-generation sequencing. OTUs from next-generation sequencing are displayed if the OTU contained more than two sequences in the unrarefied OTU table (3626 OTUs). (from Marine prokaryotes)
Solar radiation can have positive (+) or negative (−) effects resulting in increases or decreases in the heterotrophic activity of bacterioplankton. (from Marine prokaryotes)
Mycoloop links between phytoplankton and zooplankton
Chytrid‐mediated trophic links between phytoplankton and zooplankton (mycoloop). While small phytoplankton species can be grazed upon by zooplankton, large phytoplankton species constitute poorly edible or even inedible prey. Chytrid infections on large phytoplankton can induce changes in palatability, as a result of host aggregation (reduced edibility) or mechanistic fragmentation of cells or filaments (increased palatability). First, chytrid parasites extract and repack nutrients and energy from their hosts in form of readily edible zoospores. Second, infected and fragmented hosts including attached sporangia can also be ingested by grazers (i.e. concomitant predation). (from Marine fungi)
Image 104640 μm microplastic found in the deep sea amphipod Eurythenes plasticus (from Marine habitat)
Image 105Oceanic pelagic food web showing energy flow from micronekton to top predators. Line thickness is scaled to the proportion in the diet. (from Marine food web)
Image 106Some lobe-finned fishes, like the extinct Tiktaalik, developed limb-like fins that could take them onto land (from Marine vertebrate)
Image 110Whales were close to extinction until legislation was put in place. (from Marine conservation)
Image 111Cnidarians are the simplest animals with cells organised into tissues. Yet the starlet sea anemone contains the same genes as those that form the vertebrate head. (from Marine invertebrates)
Estimates of microbial species counts in the three domains of life
Bacteria are the oldest and most biodiverse group, followed by Archaea and Fungi (the most recent groups). In 1998, before awareness of the extent of microbial life had gotten underway, Robert M. May estimated there were 3 million species of living organisms on the planet. But in 2016, Locey and Lennon estimated the number of microorganism species could be as high as 1 trillion. (from Marine prokaryotes)
Image 117Dickinsonia may be the earliest animal. They appear in the fossil record 571 million to 541 million years ago. (from Marine invertebrates)
Image 118Antarctic marine food web. Potter Cove 2018. Vertical position indicates trophic level and node widths are proportional to total degree (in and out). Node colors represent functional groups. (from Marine food web)
Image 119Ernst Haeckel's 96th plate, showing some marine invertebrates. Marine invertebrates have a large variety of body plans, which are currently categorised into over 30 phyla. (from Marine invertebrates)
Image 121Waves and currents shape the intertidal shoreline, eroding the softer rocks and transporting and grading loose particles into shingles, sand or mud (from Marine habitat)
Image 122Morphological diversity of fungi collected from a marine sponge species, Ircinia variabilis (from Marine fungi)
Image 124Archaea were initially viewed as extremophiles living in harsh environments, such as the yellow archaea pictured here in a hot spring, but they have since been found in a much broader range of habitats. (from Marine prokaryotes)
Image 4Global map of large marine ecosystems. Oceanographers and biologists have identified 66 LMEs worldwide. (from Marine ecosystem)
Image 5Ecosystem services delivered by epibenthicbivalve reefs. Reefs provide coastal protection through erosion control and shoreline stabilization, and modify the physical landscape by ecosystem engineering, thereby providing habitat for species by facilitative interactions with other habitats such as tidal flat benthic communities, seagrasses and marshes. (from Marine ecosystem)
... The sea otter often keeps a stone tool in its armpit pouch.
... The ear bone called the hammer (malleus) in cetaceans is fused to the walls of the bone cavity where the ear bones are, making hearing in air nearly impossible. Instead sound is transmitted through their jaws and skull bones.
... dolphins often leap clear of the water when travelling at speed. This is because the density of water is much greater than that of air and they are able to travel faster by leaping out of the water.
... the Sperm Whale, at 18 metres long, is the largest toothed animal to have ever lived.
... If sharks don’t keep on swimming they sink to the seabed.
... some cetaceans can dive to depths of more than a kilometre and stay there for more than an hour.
The Magellanic penguin (Spheniscus magellanicus) is a South American penguin, breeding in coastal Argentina, Chile and the Falkland Islands, with some migrating to Brazil. It is the most numerous of the Spheniscus penguins. Its nearest relatives are the African Penguin, the Humboldt Penguin and the Galápagos Penguin.
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![Image 51Pennate diatom from an Arctic meltpond, infected with two chytrid-like [zoo-]sporangium fungal pathogens (in false-colour red). Scale bar = 10 μm. (from Marine fungi)](/wiki/File:Pennate_diatom_infected_with_two_chytrid-like_fungal_pathogens.png)
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